coaA Resolved · high auto-curated

H37Rv Rv1092c · MTBC0 - · 312 aa · 1219248–1220186 (-) · RefSeq NP_215608.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)pantothenate kinase
MTBC0 PGAP re-annotation
Revised (this work)Pantothenate kinase. Pfam: PRK (PF00485.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Annotated on the H37Rv protein: this gene has no 1:1 ancestral MTBC0 anchor (PE/PPE, paralogue, IS element, or otherwise unanchored CDS).

Curated reference (UniProt)

UniProt P9WPA7 SwissProt · reviewed · Evidence at protein level
UniProt namePantothenate kinase
EC (curated) EC 2.7.1.33

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category F Nucleotide transport and metabolism
Preferred namecoaA
eggNOG descriptionPantothenic acid kinase
Orthologous groupCOG0572
EC number EC 2.7.1.33
KEGG orthology K00867
KEGG pathways map00770, map01100
KEGG modules M00120
Gene Ontology (76) GO:0003674, GO:0003824, GO:0004594, GO:0005575, GO:0005622, GO:0005623, GO:0005737, GO:0005829, GO:0005886, GO:0006139, GO:0006163, GO:0006164 +64 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.231 · purifying
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
PRKPF00485.25 5.4e-1192–229 Phosphoribulokinase / Uridine kinase family

Functional interaction network (STRING v12, guilt-by-association)

PartnerProductScoreNo text-miningChannels (≥400)
Rv1391 dfp bifunctional phosphopantothenoylcysteine decarboxylase/phosphopantothenate--cysteine ligase 963 904 database:900 textmining:640
Rv3602c panC pantothenate synthetase 963 900 database:900 textmining:650
Rv2965c kdtB phosphopantetheine adenylyltransferase 959 900 database:900 textmining:614
Rv3600c coaX type III pantothenate kinase 945 900 database:900 textmining:482
Rv3433c nnr bifunctional ADP-dependent (S)-NAD(P)H-hydrate dehydratase/NAD(P)H-hydrate epimerase 411 412
Rv1093 glyA1 serine hydroxymethyltransferase 407 398
Rv3790 dprE1 decaprenylphosphoryl-beta-D-ribose oxidase 532 200 textmining:440
Rv1631 coaE dephospho-CoA kinase CoaE 692 168 textmining:645
Rv0429c def polypeptide deformylase 521 63 textmining:510
Rv0117 oxyS oxidative stress response regulatory protein OxyS 435 51 textmining:430
Rv0507 mmpL2 transmembrane transport protein MmpL2 434 47 textmining:431
Rv1971 mce3F Mce family protein Mce3F 513 41 textmining:513
Rv2225 panB 3-methyl-2-oxobutanoate hydroxymethyltransferase 481 41 textmining:482

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Annotation from H37Rv (no MTBC0 1:1 anchor; H37Rv protein used): pantothenate kinase
  • Pfam (hmmscan --cut_ga): PRK PF00485.25 (E=5e-11)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215608.1)
  • Domains: Pfam-A via hmmscan --cut_ga — PRK (PF00485.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0572
  • Curated reference: UniProt P9WPA7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 13 functional partner(s)
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>H37Rv|Rv1092c|coaA
MSRLSEPSPYVEFDRRQWRALRMSTPLALTEEELVGLRGLGEQIDLLEVEEVYLPLARLIHLQVAARQRLFAATAEFLGEPQQNPDRPVPFIIGVAGSVAVGKSTTARVLQALLARWDHHPRVDLVTTDGFLYPNAELQRRNLMHRKGFPESYNRRALMRFVTSVKSGSDYACAPVYSHLHYDIIPGAEQVVRHPDILILEGLNVLQTGPTLMVSDLFDFSLYVDARIEDIEQWYVSRFLAMRTTAFADPESHFHHYAAFSDSQAVVAAREIWRTINRPNLVENILPTRPRATLVLRKDADHSINRLRLRKL