Rv0759c Family assigned · medium auto-curated

H37Rv Rv0759c · MTBC0 - · 110 aa · 853825–854157 (-) · RefSeq NP_215273.3

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotation
Revised (this work)Contains HIT (PF01230.30) domain(s); putative function inferred from the domain architecture.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Annotated on the H37Rv protein: this gene has no 1:1 ancestral MTBC0 anchor (PE/PPE, paralogue, IS element, or otherwise unanchored CDS).

Curated reference (UniProt)

UniProt P9WML3 SwissProt · reviewed · Evidence at protein level
UniProt nameUncharacterized HIT-like protein Rv0759c

UniProt still lists this protein as Uncharacterized HIT-like protein Rv0759c; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category F Nucleotide transport and metabolism
G Carbohydrate transport and metabolism
Preferred namehit
eggNOG descriptiondiadenosine tetraphosphate (Ap4A) hydrolase and other hit family hydrolases
Orthologous groupCOG0537
KEGG orthology K02503

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.0 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 0 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
HITPF01230.30 2.2e-202–77 HIT domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: adhB (alcohol dehydrogenase B), medium confidence from genomic context alone (score 563 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv1684 hyp hypothetical protein 783 783 coexpression:775
Rv0668 rpoC DNA-directed RNA polymerase subunit beta' 656 644 database:592
Rv1390 rpoZ DNA-directed RNA polymerase subunit omega 622 622 database:585
Rv0667 rpoB DNA-directed RNA polymerase subunit beta 610 604 database:586
Rv3457c rpoA DNA-directed RNA polymerase subunit alpha 603 603 database:595
Rv3211 rhlE ATP-dependent RNA helicase RhlE 609 601 database:538
Rv1253 deaD ATP-dependent RNA helicase DeaD 607 600 database:538
Rv0861c ercc3 DNA helicase Ercc3 595 596 database:543
Rv2101 helZ helicase HelZ 573 573 database:563
Rv1329c dinG ATP-dependent helicase DinG 566 567 database:563
Rv1179c hyp hypothetical protein 565 565 database:543
Rv1947 hyp hypothetical protein 564 565 database:543
Rv0761c adhB alcohol dehydrogenase B 563 563 ctx neighborhood:530
Rv2917 hyp hypothetical protein 562 563 database:543
Rv1961 hyp hypothetical protein 561 562 database:543

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Annotation from H37Rv (no MTBC0 1:1 anchor; H37Rv protein used): hypothetical protein
  • Pfam (hmmscan --cut_ga): HIT PF01230.30 (E=2e-20)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215273.3)
  • Domains: Pfam-A via hmmscan --cut_ga — HIT (PF01230.30)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0537
  • Curated reference: UniProt P9WML3 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 43 functional partner(s); context anchor adhB
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>H37Rv|Rv0759c|
MAFLTIEPMTQGHTLVVPRAEIDHWQNVDPALFGRVMSVSQLIGKAVCRAFSTQRAGMIIAGLEVPHLHIHVFPTRSLSDFGFANVDRNPSPGSLDEAQAKIRAALAQLA