Rv0248c Family assigned · medium auto-curated

H37Rv Rv0248c · MTBC0 mtbc0_000264 · 646 aa · 299245–301185 (-) · RefSeq NP_214762.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)succinate dehydrogenase flavoprotein subunit
MTBC0 PGAP re-annotationfumarate reductase/succinate dehydrogenase flavoprotein subunit
Revised (this work)Fumarate reductase/succinate dehydrogenase flavoprotein subunit. Pfam: FAD_binding_2 (PF00890.31), Succ_DH_flav_C (PF02910.26).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53670 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable succinate dehydrogenase [iron-sulfur subunit]

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category C Energy production and conversion
Preferred namesdhA
eggNOG descriptionsuccinate dehydrogenase
Orthologous groupCOG1053
EC number EC 1.3.5.1, EC 1.3.5.4, EC 1.4.3.16
KEGG orthology K00239, K00244, K00278
KEGG pathways map00020, map00190, map00250, map00620, map00650, map00720, map00760, map01100, map01110, map01120, map01130, map01200, map02020, map05134
KEGG modules M00009, M00011, M00115, M00149, M00150, M00173, M00374, M00376
Gene Ontology (75) GO:0000104, GO:0000166, GO:0001539, GO:0003674, GO:0003824, GO:0005488, GO:0005575, GO:0005622, GO:0005623, GO:0005737, GO:0005829, GO:0005886 +63 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.194 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
FAD_binding_2PF00890.31 6.0e-8710–429 FAD binding domain
Succ_DH_flav_CPF02910.26 2.4e-21488–587 Fumarate reductase flavoprotein C-term

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0247c (succinate dehydrogenase iron-sulfur subunit), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0247c succinate dehydrogenase iron-sulfur subunit 999 1000 ctx neighborhood:881 cooccurence:774 coexpression:882 experimental:997 database:956 textmining:852
Rv3316 sdhC succinate dehydrogenase cytochrome B-556 subunit 999 996 coexpression:652 experimental:707 database:960 textmining:812
Rv0249c succinate dehydrogenase membrane anchor subunit 998 995 ctx neighborhood:839 cooccurence:771 experimental:870 textmining:655
Rv1553 frdB fumarate reductase iron-sulfur subunit 996 994 ctx cooccurence:585 coexpression:484 experimental:454 database:956 textmining:426
Rv3319 sdhB succinate dehydrogenase iron-sulphur protein subunit 997 993 ctx cooccurence:446 coexpression:489 experimental:454 database:956 textmining:628
Rv3317 sdhD succinate dehydrogenase hydrophobic membrane anchor subunit 998 992 coexpression:650 experimental:784 database:900 textmining:794
Rv0951 sucC succinyl-CoA ligase subunit beta 982 970 coexpression:584 database:900 textmining:452
Rv0952 sucD succinyl-CoA ligase subunit alpha 979 969 coexpression:588 database:900
Rv1554 frdC fumarate reductase membrane anchor subunit 975 967 coexpression:401 experimental:469 database:900
Rv1098c fum fumarate hydratase 970 963 coexpression:531 database:900
Rv1555 frdD fumarate reductase membrane anchor subunit 971 961 experimental:455 database:900
Rv1731 gabD2 succinate-semialdehyde dehydrogenase 933 930 database:900
Rv0234c gabD1 succinate-semialdehyde dehydrogenase 932 930 database:900
Rv1248c kgd multifunctional 2-oxoglutarate dehydrogenase E1 component /2-oxoglutarate dehydrogenase dihydrolipoyllysine-residue succinyltransferase 945 920 coexpression:897
Rv1552 frdA fumarate reductase flavoprotein subunit 921 918 database:900

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: succinate dehydrogenase flavoprotein subunit
  • MTBC0 PGAP product: fumarate reductase/succinate dehydrogenase flavoprotein subunit
  • Pfam (hmmscan --cut_ga): FAD_binding_2 PF00890.31 (E=6e-87), Succ_DH_flav_C PF02910.26 (E=2e-21)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214762.1)
  • Domains: Pfam-A via hmmscan --cut_ga — FAD_binding_2 (PF00890.31), Succ_DH_flav_C (PF02910.26)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1053
  • Curated reference: UniProt O53670 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 71 functional partner(s); context anchor Rv0247c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000264|Rv0248c|
MVEVERHSYDVVVIGAGGAGLRAVIEARERGLKVAVVCKSLFGKAHTVMAEGGCAAAMGNANPKDNWKTHFGDTMRGGKFLNNWRMAELHAKEAPDRVWELETYGALFDRTDDGRISQRNFGGHTYPRLAHVGDRTGLELIRTLQQKVVSLQQEDHAELGDYEARIKVFAECTITELLKDQGAIAGAFGYWRESGRFIVFEAPAVVLATGGIGKSFKVTSNSWEYTGDGHALALRAGATLINMEFVQFHPTGMVWPPSVKGILVTEGVRGDGGVLKNSENSRFMFDYIPPVFKGQYAETEEEADQWLKDNDSARRTPDLLPRDEVARAINSEVKAGRGTPHGGVYLDIASRLTPAEIKRRLPSMYHQFKELAEVDITTQAMEVGPTCHYVMGGVEVDADTGAATVPGLFAAGECAGGMHGSNRLGGNSLSDLLVFGRRAGLGAADYVRALSSRPAVSAEAIDAAAQQALSPFEGPKDGSAPENPYALHMDLQYVMNDLVGIIRNADEISRALTLLAELWSRYHNVLVEGHRQYNPGWNLSIDLRNMLLVSECVARAALQRTESRGGHTRDDHPGMDPNWRRILLVCRATETMGTGGSGSGDSNCHINVTQQLQTPMRPDLLELFEISELEKYYTDEELAEHPGRRG