Rv0249c Family assigned · low auto-curated

H37Rv Rv0249c · MTBC0 mtbc0_000265 · 273 aa · 301216–302037 (-) · RefSeq NP_214763.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)succinate dehydrogenase membrane anchor subunit
MTBC0 PGAP re-annotationsuccinate dehydrogenase membrane anchor subunit
Revised (this work)Succinate dehydrogenase membrane anchor subunit.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53671 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable succinate dehydrogenase [membrane anchor subunit]

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionmembrane
Orthologous group2CEY7

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.193 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0247c (succinate dehydrogenase iron-sulfur subunit), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0247c succinate dehydrogenase iron-sulfur subunit 999 1000 ctx neighborhood:839 cooccurence:774 experimental:997 textmining:870
Rv0248c succinate dehydrogenase flavoprotein subunit 998 995 ctx neighborhood:839 cooccurence:771 experimental:870 textmining:655
Rv0250c hyp hypothetical protein 940 879 ctx neighborhood:787 cooccurence:451 textmining:527
Rv3319 sdhB succinate dehydrogenase iron-sulphur protein subunit 905 801 experimental:784 textmining:543
Rv1553 frdB fumarate reductase iron-sulfur subunit 898 800 experimental:784 textmining:511
Rv3318 sdhA succinate dehydrogenase flavoprotein subunit 657 512 experimental:469
Rv1552 frdA fumarate reductase flavoprotein subunit 634 509 experimental:469
Rv3687c rsfB anti-anti-sigma factor RsfB 478 478 ctx cooccurence:476
Rv3537 kstD 3-oxosteroid 1-dehydrogenase 409 410
Rv1817 flavoprotein 409 410
Rv3818 hyp hypothetical protein 401 402
Rv0251c hsp heat shock protein 531 399
Rv3316 sdhC succinate dehydrogenase cytochrome B-556 subunit 805 53 textmining:803
Rv3317 sdhD succinate dehydrogenase hydrophobic membrane anchor subunit 802 49 textmining:801
Rv0392c ndhA NADH dehydrogenase NdhA 511 41 textmining:511

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: succinate dehydrogenase membrane anchor subunit
  • MTBC0 PGAP product: succinate dehydrogenase membrane anchor subunit
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214763.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2CEY7
  • Curated reference: UniProt O53671 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 15 functional partner(s); context anchor Rv0247c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000265|Rv0249c|
MSAPTANRPAIGVFTPTRAQIPERTLRTDLWWLPPLLTNLGLLAFICYATTRAFWGSQYWVEKYHYLTPFYSPCVSASCQPGASHLGVWFGHFPGWIPLGAMVLPFLLGFRLTCYYYRKAYYRSVWQSPTSCAVPEPRAHYTGETRLPLIVQNTHRYFFYIAVVVSLINTYDAIAAFHSPSGFGFGLGNVILTINVVLLWAYTISCHSCRHATGGRLKHFSKHPVRYWIWTQVSKLNTRHMQFAWITLGTLALTDFYIMLVASGSITDLRFIG