fadD7 Family assigned · medium auto-curated

H37Rv Rv0119 · MTBC0 mtbc0_000130 · 525 aa · 144215–145792 (+) · RefSeq NP_214633.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)fatty-acid--CoA ligase FadD7
MTBC0 PGAP re-annotationFadD7 family fatty acid--CoA ligase
Revised (this work)FadD7 family fatty acid--CoA ligase. Pfam: AMP-binding (PF00501.35), AMP-binding_C (PF13193.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O07169 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable fatty-acid-CoA ligase FadD7

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category I Lipid transport and metabolism
Q Secondary metabolites biosynthesis, transport and catabolism
Preferred namefadD7
eggNOG descriptionAMP-binding enzyme C-terminal domain
Orthologous groupCOG0318
KEGG orthology K16029
KEGG pathways map01051, map01052

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.241 · purifying
Polymorphic sites (≥ 0.1% of strains) 11 synonymous, 7 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
AMP-bindingPF00501.35 9.0e-7314–386 AMP-binding enzyme
AMP-binding_CPF13193.13 1.5e-17437–510 AMP-binding enzyme C-terminal domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: oxcA (oxalyl-CoA decarboxylase OxcA), high confidence from genomic context alone (score 893 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0118c oxcA oxalyl-CoA decarboxylase OxcA 895 893 ctx neighborhood:720 coexpression:454
Rv0719 rplF 50S ribosomal protein L6 694 694 experimental:402 database:510
Rv2947c pks15 polyketide synthase 682 682 ctx fusion:452
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 706 681
Rv1527c pks5 polyketide synthase 705 680
Rv2048c pks12 polyketide synthase 705 680
Rv2933 ppsC phthiocerol synthesis polyketide synthase type I PpsC 704 679
Rv2940c mas multifunctional mycocerosic acid synthase 704 679
Rv1180 pks3 polyketide beta-ketoacyl synthase 662 662 ctx fusion:414
Rv2382c mbtC polyketide synthetase 640 637
Rv3800c pks13 polyketide synthase 669 636
Rv2946c pks1 polyketide synthase 663 631
Rv2932 ppsB phthiocerol synthesis polyketide synthase type I PpsB 651 631
Rv0405 pks6 membrane bound polyketide synthase 645 628
Rv2380c mbtE peptide synthetase 635 619 experimental:465

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: fatty-acid--CoA ligase FadD7
  • MTBC0 PGAP product: FadD7 family fatty acid--CoA ligase
  • Pfam (hmmscan --cut_ga): AMP-binding PF00501.35 (E=9e-73), AMP-binding_C PF13193.13 (E=2e-17)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214633.1)
  • Domains: Pfam-A via hmmscan --cut_ga — AMP-binding (PF00501.35), AMP-binding_C (PF13193.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0318
  • Curated reference: UniProt O07169 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 69 functional partner(s); context anchor oxcA
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000130|Rv0119|fadD7
MASDFGPRIADLVEVAATRLPEAPALVVTADRIAISHRDLARLVDELAGQLTRSGLLPGDRVALRMGSNAEFVVALLAASRADLVVVPLDPALPITEQRVRSQAAGARVVLIDADGPHDRAEPTTRWWPLTVNVGGDSGPSGGTLSVHLDAATEPNPATSTPEGLRPDDAMIMFTGGTTGLPKMVPWTHANIASSVRAIITGYRLSPRDATVAVMPLYHGHGLIASLLATLASGGAVSLPARGRFSAHTFWDDIKAVGATWYTAVPTIHQILLERSATEPSGRKPAALRFIRSCSAPLTAQAALALQTEFAAPVVCAFGMTEATHQVTTTQIEGIDQTETPVVSTGLVGRSTGAQIRIVGSDGLPLPAGAVGEIWLRGTTVVRGYLGDPTITAANFTDGWLRTGDLGSLSAAGDLSIRGRIKELINRGGEKISPERVEGVLASHPNVMEAAVFGVPHQLYGEAVAAVIVPRESAPPTREELVQFCRERLAAFEIPASFQEASGLPHTAKGSLDRRAVAERFGHSV