Rv3714c Family assigned · medium auto-curated

H37Rv Rv3714c · MTBC0 mtbc0_003937 · 296 aa · 4183052–4183942 (-) · RefSeq NP_218231.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationhypothetical protein
Revised (this work)Contains AbiEi_1 (PF09407.17) domain(s); putative function inferred from the domain architecture.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O69681 TrEMBL · unreviewed · Evidence at protein level
UniProt nameCullin, a subunit of E3 ubiquitin ligase

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category K Transcription
eggNOG descriptionPsort location Cytoplasmic, score
Orthologous groupCOG5340

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.637 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 9 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
AbiEi_1PF09407.17 1.7e-0578–185 AbiEi antitoxin C-terminal domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: recR (recombination protein RecR), medium confidence from genomic context alone (score 636 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv3585 radA DNA repair protein RadA 791 791 coexpression:791
Rv3586 disA DNA integrity scanning protein DisA 730 730 coexpression:730
Rv3715c recR recombination protein RecR 635 636 ctx neighborhood:632
Rv3716c hyp hypothetical protein 627 627 ctx neighborhood:625
Rv3859c gltB glutamate synthase large subunit 522 522 coexpression:515
Rv3776 hyp hypothetical protein 478 479 ctx cooccurence:451
Rv2100 hyp hypothetical protein 473 474 ctx cooccurence:447
Rv0393 hyp hypothetical protein 471 471 ctx cooccurence:467
Rv1702c hyp hypothetical protein 448 448 ctx cooccurence:412
Rv0515 hyp hypothetical protein 446 447
Rv0767c HTH-type transcriptional regulator 445 445 ctx cooccurence:443
Rv3810 pirG cell surface protein 442 443 ctx cooccurence:440
Rv3717 hyp hypothetical protein 431 432 ctx neighborhood:425
Rv3497c mce4C Mce family protein Mce4C 430 261
Rv0290 eccD3 ESX-3 secretion system protein EccD 525 205 textmining:428

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: hypothetical protein
  • Pfam (hmmscan --cut_ga): AbiEi_1 PF09407.17 (E=2e-05)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218231.1)
  • Domains: Pfam-A via hmmscan --cut_ga — AbiEi_1 (PF09407.17)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG5340
  • Curated reference: UniProt O69681 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 21 functional partner(s); context anchor recR
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003937|Rv3714c|
MLISRMSVRSASMSVMGDVFIGSEAITAGRLTRHELQRWYQPMFRGVYVSRRSVPTLWDRTVGAWLATRRHGVIAGNAASALHGAQWVDVDVAIELISPTTRPQHGLVIRRETLCDDEITRVVGLPVTTLARTAYDLGRHLSRGEAVARLDALMRATPFSRDDVLLLAKRHAGARGVRRLRDVLPLVDGGAASPKETWLRLLLIDAGLPVPTTQIPVVHRWRNVGVLDMGWEKYMVAAEYDGDQHRSDRGRYVKDQRRLRKLAELGWIVIRVIAEDNPDDVVNRVRAALLARGWRP