Rv3680 Family assigned · medium auto-curated

H37Rv Rv3680 · MTBC0 mtbc0_003900 · 393 aa · 4143625–4144806 (+) · RefSeq NP_218197.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)anion transporter ATPase
MTBC0 PGAP re-annotationArsA family ATPase
Revised (this work)ArsA family ATPase. Pfam: ArsA_ATPase (PF02374.22), CbiA (PF01656.30), AAA_31 (PF13614.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6Y498 SwissProt · reviewed · Evidence at protein level
UniProt nameATPase BagB
EC (curated) EC 3.6.1.-
Curated functionComponent of the heterodimeric BagAB ATPase complex, whose two subunits are actively involved in ATP hydrolysis. The ATPase activity is required to mediate resistance against nitric oxide (NO) and elevated levels of glycerol.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category D Cell cycle control, cell division, chromosome partitioning
eggNOG descriptionanion-transporting ATPase
Orthologous groupCOG0003

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.715 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 6 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
ArsA_ATPasePF02374.22 1.2e-2520–285 Anion-transporting ATPase
CbiAPF01656.30 4.7e-0821–259 CobQ/CobB/MinD/ParA nucleotide binding domain
AAA_31PF13614.13 1.1e-0926–162 AAA domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3679 (anion transporter ATPase), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3679 anion transporter ATPase 999 1000 ctx neighborhood:882 experimental:999 database:900
Rv3678A hyp hypothetical protein 785 785 ctx neighborhood:783
Rv3677c beta lactamase 788 781 ctx neighborhood:774
Rv3678c hyp hypothetical protein 777 777 ctx neighborhood:774
Rv0487 hyp hypothetical protein 712 713 ctx cooccurence:709
Rv0502 hyp hypothetical protein 668 668 ctx cooccurence:653
Rv2185c TB16.3 hyp hypothetical protein 650 651 ctx cooccurence:635
Rv0244c fadE5 acyl-CoA dehydrogenase FadE5 638 638 ctx cooccurence:635
Rv1467c fadE15 acyl-CoA dehydrogenase 612 613 ctx cooccurence:607
Rv0854 hyp hypothetical protein 605 605 ctx cooccurence:591
Rv0857 hyp hypothetical protein 584 585 ctx cooccurence:582
Rv2267c stf3 hyp hypothetical protein 578 557 database:410
Rv1691 hyp hypothetical protein 578 557 database:410
Rv3529c hyp hypothetical protein 577 556 database:410
Rv0501 galE2 UDP-glucose 4-epimerase GalE 555 555 ctx cooccurence:540

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: anion transporter ATPase
  • MTBC0 PGAP product: ArsA family ATPase
  • Pfam (hmmscan --cut_ga): ArsA_ATPase PF02374.22 (E=1e-25), CbiA PF01656.30 (E=5e-08), AAA_31 PF13614.13 (E=1e-09)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218197.1)
  • Domains: Pfam-A via hmmscan --cut_ga — ArsA_ATPase (PF02374.22), CbiA (PF01656.30), AAA_31 (PF13614.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0003
  • Curated reference: UniProt I6Y498 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 37 functional partner(s); context anchor Rv3679
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003900|Rv3680|
MSVTPKTLDMGAILADTSNRVVVCCGAGGVGKTTTAAALALRAAEYGRTVVVLTIDPAKRLAQALGINDLGNTPQRVPLAPEVPGELHAMMLDMRRTFDEMVMQYSGPERAQSILDNQFYQTVATSLAGTQEYMAMEKLGQLLSQDRWDLIVVDTPPSRNALDFLDAPKRLGSFMDSRLWRLLLAPGRGIGRLITGVMGLAMKALSTVLGSQMLADAAAFVQSLDATFGGFREKADRTYALLKRRGTQFVVVSAAEPDALREASFFVDRLSQESMPLAGLVFNRTHPMLCALPIERAIDAAETLDAETTDSDATSLAAAVLRIHAERGQTAKREIRLLSRFTGANPTVPVVGVPSLPFDVSDLEALRALADQLTTVGNDAGRAAPAAEEPAHQ