Rv3552 Family assigned · low auto-curated

H37Rv Rv3552 · MTBC0 mtbc0_003769 · 250 aa · 4014448–4015200 (+) · RefSeq NP_218069.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)CoA-transferase subunit beta
MTBC0 PGAP re-annotationCoA-transferase subunit beta
Revised (this work)CoA-transferase subunit beta.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WPV9 SwissProt · reviewed · Evidence at protein level
UniProt nameCholesterol ring-cleaving hydrolase IpdB subunit
EC (curated) EC 4.1.99.-
Curated functionInvolved in the final steps of cholesterol and steroid degradation. Opens the last steroid ring of cholesterol by catalyzing the hydrolysis of (3E)-2-(2-carboxylatoethyl)-3-methyl-6-oxocyclohex-1-ene-1-carboxyl-CoA (COCHEA-CoA) to 6-methyl-3,7-dioxodecanedioyl-CoA (MeDODA-CoA).

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category I Lipid transport and metabolism
Preferred namecatJ
eggNOG descriptionCOG2057 Acyl CoA acetate 3-ketoacid CoA transferase, beta subunit
Orthologous groupCOG2057
Gene Ontology (8) GO:0008150, GO:0040007, GO:0044110, GO:0044116, GO:0044117, GO:0044403, GO:0044419, GO:0051704

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS n/a
Polymorphic sites (≥ 0.1% of strains) 0 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3551 (CoA-transferase subunit alpha), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3551 CoA-transferase subunit alpha 999 1000 ctx neighborhood:882 cooccurence:774 coexpression:952 experimental:999 database:900 textmining:820
Rv3550 echA20 enoyl-CoA hydratase EchA20 999 998 ctx neighborhood:882 cooccurence:749 coexpression:869 database:500 textmining:564
Rv3549c short-chain type dehydrogenase/reductase 984 983 ctx neighborhood:787 cooccurence:593 coexpression:807
Rv3553 oxidoreductase 980 981 ctx neighborhood:773 cooccurence:446 coexpression:860
Rv3548c short-chain type dehydrogenase/reductase 957 956 ctx neighborhood:588 cooccurence:455 coexpression:800
Rv3559c oxidoreductase 943 941 ctx cooccurence:709 coexpression:783
Rv3560c fadE30 acyl-CoA dehydrogenase FadE30 964 922 ctx cooccurence:668 coexpression:771 textmining:564
Rv2504c scoA succinyl-CoA:3-ketoacid-CoA transferase subunit A 928 922 coexpression:670 experimental:766
Rv3556c fadA6 acetyl-CoA acetyltransferase FadA 966 904 coexpression:780 database:500 textmining:667
Rv3562 fadE31 acyl-CoA dehydrogenase FadE31 904 901 ctx cooccurence:736 coexpression:631
Rv3561 fadD3 fatty-acid--CoA ligase FadD3 881 877 coexpression:810
Rv3522 ltp4 lipid transfer protein 824 816 ctx cooccurence:761
Rv3540c ltp2 lipid transfer protein 813 805 ctx cooccurence:750
Rv3523 ltp3 lipid carrier protein 799 791 ctx cooccurence:732
Rv3541c chsH1 hyp hypothetical protein 791 784 ctx cooccurence:769

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: CoA-transferase subunit beta
  • MTBC0 PGAP product: CoA-transferase subunit beta
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218069.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2057
  • Curated reference: UniProt P9WPV9 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 57 functional partner(s); context anchor Rv3551
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003769|Rv3552|
MSTRAEVCAVACAELFRDAGEIMISPMTNMASVGARLARLTFAPDILLTDGEAQLLADTPALGKTGAPNRIEGWMPFGRVFETLAWGRRHVVMGANQVDRYGNQNISAFGPLQRPTRQMFGVRGSPGNTINHATSYWVGNHCKRVFVEAVDVVSGIGYDKVDPDNPAFRFVNVYRVVSNLGVFDFGGPDHSMRAVSLHPGVTPGDVRDATSFEVHDLDAAEQTRLPTDDELHLIRAVIDPKSLRDREIRS