pgsA3 Resolved · high auto-curated

H37Rv Rv2746c · MTBC0 - · 209 aa · 3058602–3059231 (-) · RefSeq NP_217262.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)CDP-diacylglycerol--glycerol-3-phosphate 3-phosphatidyltransferase
MTBC0 PGAP re-annotation
Revised (this work)CDP-diacylglycerol--glycerol-3-phosphate 3-phosphatidyltransferase. Pfam: CDP-OH_P_transf (PF01066.27).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Annotated on the H37Rv protein: this gene has no 1:1 ancestral MTBC0 anchor (PE/PPE, paralogue, IS element, or otherwise unanchored CDS).

Curated reference (UniProt)

UniProt P9WPG3 SwissProt · reviewed · Inferred from homology
UniProt nameProbable phosphatidylglycerophosphate synthase
EC (curated) EC 2.7.8.5
Curated functionProbably catalyzes the synthesis of phosphatidylglycerophosphate by transferring a phosphatidyl group from CDP-diacylglycerol to glycerol 3-phosphate.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category I Lipid transport and metabolism
Preferred namepgsA
eggNOG descriptionBelongs to the CDP-alcohol phosphatidyltransferase class-I family
Orthologous groupCOG0558
EC number EC 2.7.8.41, EC 2.7.8.5
KEGG orthology K00995, K08744
KEGG pathways map00564, map01100
Gene Ontology (36) GO:0003674, GO:0003824, GO:0005575, GO:0005618, GO:0005623, GO:0006629, GO:0006644, GO:0006650, GO:0006793, GO:0006796, GO:0008150, GO:0008152 +24 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.373 · purifying
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
CDP-OH_P_transfPF01066.27 3.9e-4628–187 CDP-alcohol phosphatidyltransferase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: argA (L-glutamate alpha-N-acetyltranferase), high confidence from genomic context alone (score 721 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1822 pgsA2 CDP-diacylglycerol--glycerol-3-phosphate 3-phosphatidyltransferase 933 928 database:900
Rv2881c cdsA phosphatidate cytidylyltransferase 964 919 database:900 textmining:574
Rv0436c pssA CDP-diacylglycerol--serine O-phosphatidyltransferase 985 918 database:900 textmining:825
Rv2612c pgsA1 CDP-diacylglycerol--inositol 3-phosphatidyltransferase 987 900 database:900 textmining:876
Rv2289 cdh CDP-diacylglycerol pyrophosphatase 900 900 database:900
Rv1901 cinA competence damage-inducible protein CinA 880 843 coexpression:733
Rv2747 argA L-glutamate alpha-N-acetyltranferase 721 721 ctx neighborhood:649
Rv2745c clgR transcriptional regulator ClgR 658 658 ctx neighborhood:656
Rv2744c 35kd_ag hyp hypothetical protein 522 503 ctx neighborhood:503
Rv2743c hyp hypothetical protein 487 487 ctx neighborhood:487
Rv2982c gpdA2 glycerol-3-phosphate dehydrogenase 586 442 ctx cooccurence:423
Rv3356c folD bifunctional methylenetetrahydrofolate dehydrogenase/methenyltetrahydrofolate cyclohydrolase 492 438 coexpression:417
Rv3279c birA bifunctional biotin operon repressor/biotin--[acetyl-CoA-carboxylase 449 429 coexpression:413
Rv0564c gpdA1 glycerol-3-phosphate dehydrogenase 562 410
Rv2483c plsC bifunctional L-3-phosphoserine phosphatase/1-acyl-sn-glycerol-3-phosphate acyltransferase 419 363

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Annotation from H37Rv (no MTBC0 1:1 anchor; H37Rv protein used): CDP-diacylglycerol--glycerol-3-phosphate 3-phosphatidyltransferase
  • Pfam (hmmscan --cut_ga): CDP-OH_P_transf PF01066.27 (E=4e-46)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217262.1)
  • Domains: Pfam-A via hmmscan --cut_ga — CDP-OH_P_transf (PF01066.27)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0558
  • Curated reference: UniProt P9WPG3 (SwissProt, reviewed; Inferred from homology)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 30 functional partner(s); context anchor argA
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>H37Rv|Rv2746c|pgsA3
MSRSTRYSVAVSAQPETGQIAGRARIANLANILTLLRLVMVPVFLLALFYGGGHHSAARVVAWAIFATACITDRFDGLLARNYGMATEFGAFVDPIADKTLIGSALIGLSMLGDLPWWVTVLILTRELGVTVLRLAVIRRGVIPASWGGKLKTFVQAVAIGLFVLPLSGPLHVAAVVVMAAAILLTVITGVDYVARALRDIGGIRQTAS