cysT Family assigned · medium auto-curated

H37Rv Rv2399c · MTBC0 mtbc0_002554 · 283 aa · 2720043–2720894 (-) · RefSeq NP_216915.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)sulfate ABC transporter permease CysT
MTBC0 PGAP re-annotationsulfate ABC transporter permease subunit CysT
Revised (this work)Sulfate ABC transporter permease subunit CysT. Pfam: BPD_transp_1 (PF00528.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P71745 TrEMBL · unreviewed · Inferred from homology
UniProt nameSulfate transport system permease protein CysT
Curated functionPart of the ABC transporter complex (TC 3.A.1.6.1) involved in sulfate/thiosulfate import..; FUNCTION: Part of the ABC transporter complex CysAWTP (TC 3.A.1.6.1) involved in sulfate/thiosulfate import. Probably responsible for the translocation of the substrate across the membrane.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category O Post-translational modification, protein turnover, chaperones
Preferred namecysT
eggNOG descriptionsulfate ABC transporter
Orthologous groupCOG0555
KEGG orthology K02046
KEGG pathways map00920, map02010
KEGG modules M00185

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.382 · purifying
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 1 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
BPD_transp_1PF00528.28 1.5e-1987–275 Binding-protein-dependent transport system inner membrane component

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: cysA1 (sulfate ABC transporter ATP-binding protein CysA), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2397c cysA1 sulfate ABC transporter ATP-binding protein CysA 999 1000 ctx neighborhood:865 cooccurence:771 coexpression:952 database:900 textmining:885
Rv2400c subI sulfate ABC transporter substrate-binding lipoprotein SubI 999 1000 ctx neighborhood:881 cooccurence:774 coexpression:963 database:900 textmining:741
Rv1286 cysC adenylyl-sulfate kinase 999 1000 coexpression:999 database:900 textmining:487
Rv2398c cysW sulfate ABC transporter permease CysW 999 1000 ctx neighborhood:882 coexpression:976 database:900
Rv1285 cysD sulfate adenylyltransferase subunit 2 974 972 coexpression:675 database:900
Rv1857 modA molybdate ABC transporter substrate-binding lipoprotein ModA 969 964 ctx cooccurence:623 experimental:895
Rv1859 modC molybdenum ABC transporter ATP-binding protein ModC 830 808 coexpression:725
Rv2392 cysH phosphoadenosine phosphosulfate reductase 759 730 coexpression:677
Rv2391 sirA sulfite reductase 673 630 coexpression:609
Rv2064 cobG precorrin-3B synthase 670 626 coexpression:605
Rv2401 hyp hypothetical protein 583 582 ctx neighborhood:578
Rv3327 transposase fusion protein 409 409 coexpression:409
Rv2429 ahpD alkyl hydroperoxide reductase AphD 406 407 coexpression:407
Rv0464c hyp hypothetical protein 406 407 coexpression:407
Rv1531 hyp hypothetical protein 403 404 coexpression:404

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: sulfate ABC transporter permease CysT
  • MTBC0 PGAP product: sulfate ABC transporter permease subunit CysT
  • Pfam (hmmscan --cut_ga): BPD_transp_1 PF00528.28 (E=1e-19)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216915.1)
  • Domains: Pfam-A via hmmscan --cut_ga — BPD_transp_1 (PF00528.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0555
  • Curated reference: UniProt P71745 (TrEMBL, unreviewed; Inferred from homology)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 20 functional partner(s); context anchor cysA1
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002554|Rv2399c|cysT
MTESLVGERRAPQFRARLSGPAGPPSVRVGMAVVWLSVIVLLPLAAIVWQAAGGGWRAFWLAVSSHAAMESFRVTLTISTAVTVINLVFGLLIAWVLVRDDFAGKRIVDAIIDLPFALPTIVASLVMLALYGNNSPVGLHFQHTATGVGVALAFVTLPFVVRAVQPVLLEIDRETEEAAASLGANGAKIFTSVVLPSLTPALLSGAGLAFSRAIGEFGSVVLIGGAVPGKTEVSSQWIRTLIENDDRTGAAAISVVLLSISFIVLLILRVVGARAAKREEMAA