Rv1544 Family assigned · medium auto-curated

H37Rv Rv1544 · MTBC0 mtbc0_001651 · 267 aa · 1755903–1756706 (+) · RefSeq NP_216060.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)ketoacyl reductase
MTBC0 PGAP re-annotationSDR family oxidoreductase
Revised (this work)SDR family oxidoreductase. Pfam: adh_short (PF00106.32), KR (PF08659.17), adh_short_C2 (PF13561.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt Q10782 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible ketoacyl reductase

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionBelongs to the short-chain dehydrogenases reductases (SDR) family
Orthologous groupCOG0300
KEGG orthology K07124

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.133 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 8 synonymous, 3 missense, 0 nonsense, 1 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 0.41% of strains (590) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
adh_shortPF00106.32 7.7e-4111–196 short chain dehydrogenase
KRPF08659.17 8.4e-1111–180 KR domain
adh_short_C2PF13561.13 1.2e-2416–220 Enoyl-(Acyl carrier protein) reductase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1543 (oxidoreductase), high confidence from genomic context alone (score 967 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1543 oxidoreductase 974 967 ctx neighborhood:801 coexpression:813
Rv1546 hyp hypothetical protein 791 791 ctx neighborhood:790
Rv1545 hyp hypothetical protein 748 748 ctx neighborhood:739
Rv1547 dnaE1 DNA polymerase III subunit alpha 681 681 ctx neighborhood:678
Rv1714 oxidoreductase 600 591 ctx cooccurence:532
Rv1350 fabG2 3-oxoacyl-ACP reductase FabG 567 551 ctx cooccurence:490
Rv3549c short-chain type dehydrogenase/reductase 565 549 ctx cooccurence:488
Rv1542c glbN hemoglobin GlbN 540 540 ctx neighborhood:540
Rv1483 fabG1 3-oxoacyl-ACP reductase FabG 507 489 ctx cooccurence:414
Rv0687 NAD-dependent oxidoreductase 497 479 ctx cooccurence:408
Rv2002 fabG3 3-alpha(or 20-beta)-hydroxysteroid dehydrogenase 486 467
Rv3559c oxidoreductase 466 447
Rv2750 dehydrogenase 450 430
Rv1541c lprI lipoprotein LprI 408 408 ctx neighborhood:406
Rv3480c diacyglycerol O-acyltransferase 814 95 textmining:803

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: ketoacyl reductase
  • MTBC0 PGAP product: SDR family oxidoreductase
  • Pfam (hmmscan --cut_ga): adh_short PF00106.32 (E=8e-41), KR PF08659.17 (E=8e-11), adh_short_C2 PF13561.13 (E=1e-24)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216060.1)
  • Domains: Pfam-A via hmmscan --cut_ga — adh_short (PF00106.32), KR (PF08659.17), adh_short_C2 (PF13561.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0300
  • Curated reference: UniProt Q10782 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 27 functional partner(s); context anchor Rv1543
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001651|Rv1544|
MSLPKPNNQTTVVITGASSGIGVELARGLAGRGFPLMLVARRRERLDELADQLRQEHCVGVEVLPLDLADTQARAQLADRLRSDAIAGLCNSAGFGTSGRFWELPFARESEEVVLNALALMELTHAALPGMVKRGAGAVLNIASIAGFQPIPYMAVYSATKAFVLTFSEAVQEELHGTGVSVTALCPGPVPTEWAEIASAERFSIPLAQVSPHDVAEAAIAGMLSGKRTVVPGIVPKFVSTSGRFAPRSLLLPAIRIGNRLRGGPSR