Rv1533 Family assigned · medium auto-curated

H37Rv Rv1533 · MTBC0 mtbc0_001640 · 375 aa · 1743419–1744546 (+) · RefSeq NP_216049.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)monooxygenase
MTBC0 PGAP re-annotationnitronate monooxygenase family protein
Revised (this work)Nitronate monooxygenase family protein. Pfam: NMO (PF03060.22), FMN_dh (PF01070.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O06179 SwissProt · reviewed · Evidence at protein level
UniProt namePutative monooxygenase Rv1533
EC (curated) EC 1.13.12.-

UniProt still lists this protein as Putative monooxygenase Rv1533; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG description2-nitropropane dioxygenase
Orthologous groupCOG2070

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.602 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
NMOPF03060.22 1.9e-5523–361 Nitronate monooxygenase
FMN_dhPF01070.25 1.4e-09172–250 FMN-dependent dehydrogenase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1534 (transcriptional regulator), high confidence from genomic context alone (score 981 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1534 transcriptional regulator 980 981 ctx neighborhood:882 coexpression:843
Rv2524c fas fatty acid synthase 782 749 coexpression:713
Rv1532c hyp hypothetical protein 614 615 ctx neighborhood:609
Rv2940c mas multifunctional mycocerosic acid synthase 577 505
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 576 503
Rv2048c pks12 polyketide synthase 574 502
Rv1527c pks5 polyketide synthase 574 502
Rv2933 ppsC phthiocerol synthesis polyketide synthase type I PpsC 572 499
Rv2246 kasB 3-oxoacyl-ACP synthase 2 464 436 coexpression:418
Rv3800c pks13 polyketide synthase 567 435
Rv2245 kasA 3-oxoacyl-ACP synthase 1 463 435 coexpression:417
Rv2243 fabD malonyl CoA-acyl carrier protein transacylase 457 426 coexpression:407
Rv2946c pks1 polyketide synthase 499 424
Rv1661 pks7 polyketide synthase 478 419
Rv1181 pks4 polyketide beta-ketoacyl synthase 477 417

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: monooxygenase
  • MTBC0 PGAP product: nitronate monooxygenase family protein
  • Pfam (hmmscan --cut_ga): NMO PF03060.22 (E=2e-55), FMN_dh PF01070.25 (E=1e-09)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216049.1)
  • Domains: Pfam-A via hmmscan --cut_ga — NMO (PF03060.22), FMN_dh (PF01070.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2070
  • Curated reference: UniProt O06179 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 19 functional partner(s); context anchor Rv1534
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001640|Rv1533|
MRTRVAELLGAEFPICAFSHCRDVVAAVSNAGGFGILGAVAHSPKRLESELTWIEEHTGGKPYGVDVLLPPKYIGAEQGGIDAQQARELIPEGHRTFVDDLLVRYGIPAVTDRQRSSSAGGLHISPKGYQPLLDVAFAHDIRLIASALGPPPPDLVERAHNHDVLVAALAGTAQHARRHAAAGVDLIVAQGTEAGGHTGEVATMVLVPEVVDAVSPTPVLAAGGIARGRQIAAALALGAEGVWCGSVWLTTEEAETPPVVKDKFLAATSSDTVRSRSLTGKPARMLRTAWTDEWDRPDSPDPLGMPLQSALVSDPQLRINQAAGQPGAKARELATYFVGQVVGSLDRVRSARSVVLDMVEEFIDTVGQLQGLVQR