Rv0585c Family assigned · medium auto-curated

H37Rv Rv0585c · MTBC0 mtbc0_000615 · 795 aa · 685436–687823 (-) · RefSeq NP_215099.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)integral membrane protein
MTBC0 PGAP re-annotationlysylphosphatidylglycerol synthase transmembrane domain-containing protein
Revised (this work)Lysylphosphatidylglycerol synthase transmembrane domain-containing protein. Pfam: LPG_synthase_TM (PF03706.19).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53781 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved integral membrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category T Signal transduction mechanisms
eggNOG descriptionintegral membrane protein
Orthologous groupCOG0392
KEGG orthology K07027

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.32 · purifying
Polymorphic sites (≥ 0.1% of strains) 13 synonymous, 11 missense, 0 nonsense, 2 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 1.08% of strains (1574) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
LPG_synthase_TMPF03706.19 6.9e-27498–782 Lysylphosphatidylglycerol synthase TM region

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: mce2C (Mce family protein Mce2C), medium confidence from genomic context alone (score 613 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0591 mce2C Mce family protein Mce2C 613 613 ctx neighborhood:545
Rv0594 mce2F Mce family protein Mce2F 603 603 ctx neighborhood:537
Rv0592 mce2D Mce family protein Mce2D 600 600 ctx neighborhood:537
Rv0479c membrane protein 566 566 ctx cooccurence:566
Rv0586 mce2R HTH-type transcriptional regulator Mce2R 566 545 ctx neighborhood:545
Rv0590 mce2B Mce-family protein Mce2B; Rv0590, (MTCY19H5.32c), len: 275 aa. Mce2B; belongs to 24-membered Mycobacterium tuberculosis Mce protein family ( 545 545 ctx neighborhood:544
Rv0587 yrbE2A hyp hypothetical protein 545 545 ctx neighborhood:545
Rv0589 mce2A Mce family protein Mce2A 545 545 ctx neighborhood:544
Rv0588 yrbE2B hyp hypothetical protein 545 545 ctx neighborhood:545
Rv0590A Mce-family related protein; Rv0590A, len: 84 aa. Probable continuation of mce2B|Rv0590. Can find no frameshift to account for this. Possible 545 544 ctx neighborhood:544
Rv0593 lprL Mce family lipoprotein LprL 542 542 ctx neighborhood:537
Rv1863c integral membrane protein 535 535 ctx cooccurence:525
Rv2876 transmembrane protein 503 503 ctx cooccurence:503
Rv2180c integral membrane protein 492 492 ctx cooccurence:489
Rv1203c hyp hypothetical protein 470 470 ctx cooccurence:467

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: integral membrane protein
  • MTBC0 PGAP product: lysylphosphatidylglycerol synthase transmembrane domain-containing protein
  • Pfam (hmmscan --cut_ga): LPG_synthase_TM PF03706.19 (E=7e-27)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215099.1)
  • Domains: Pfam-A via hmmscan --cut_ga — LPG_synthase_TM (PF03706.19)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0392
  • Curated reference: UniProt O53781 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 30 functional partner(s); context anchor mce2C
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000615|Rv0585c|
MRVDGRDIGVSGNLLQPLTRRTNDIIRAVLAAIYLVAVITSSLITRPQWVALEKSISEIVGVLSPSQSDLVYLGYGLAILALPFVILIGLIVSRQWKLLGAYAAAGLMAVLPLSISSSRIAAPRWHFDLSDRLATLLAQFLDDPRWIAMLAAVLTVSGPWLPARWRHWWWALLLAFVPIHLVVSAIVPARSLLGLAVGWLVGALVVLVVGTPALEVPLDGAIRALAKRGFAVSGLAVVRPAGPGPLVLSAACEQPNAGACSEALIELYGPHQSGGGALRQLWLKLTLRGTETAPLQASMRRAVEHRALMAIAFGDLGMANTTVIAVSPLDRGWTLYAHRPARGIGISECTKTTPTAHVWEALRTLHDQQISHGDLCSAEITVDNGAVLFGGFGEAEYGATDAQLQSDLAQLLVTTSALYDAEAAVTAAIDTFGKQAILAASRRLTKSAVPKRIRESITDPNAVIASTRAEVMRQTGADQIKAETITRFSRGQLIQLVLIGALVYVAYPFISTVPTFFSQLRTANWWWALLGLAVSALTYVGAAAALWACADGLVGFWKLSIMQVANTFAATTTPAGVGGLALSTRFLQKGGLTAVRATAAVALQQSVQVIVHLVLLILFSALAGTSTDLSHFVPNATVLYLIAGVALGIVGTFLFVPKLRRWLATAVRPKLREVTNDLIALAREPKRLALIVLGCAGTTLGAALALWASIEAFGGGTTFVTVTVVTMVGGTLASAAPTPGGVGAVEAALIGGLAAFGVPAALGVPSVLLYRLLTCWLPVFAGWQVMHWLTRHEMI