Rv0584 Family assigned · medium auto-curated

H37Rv Rv0584 · MTBC0 mtbc0_000614 · 877 aa · 682780–685413 (+) · RefSeq NP_215098.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)glycosidase
MTBC0 PGAP re-annotationGH92 family glycosyl hydrolase
Revised (this work)GH92 family glycosyl hydrolase. Pfam: Glyco_hydro_92N (PF17678.8), Glyco_hydro_92 (PF07971.19).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O86365 SwissProt · reviewed · Inferred from homology
UniProt nameUncharacterized glycosidase Rv0584
EC (curated) EC 3.2.1.-

UniProt still lists this protein as Uncharacterized glycosidase Rv0584; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category G Carbohydrate transport and metabolism
eggNOG descriptionGlycosyl hydrolase family 92
Orthologous groupCOG3537
Gene Ontology (45) GO:0000224, GO:0003674, GO:0003824, GO:0005575, GO:0005622, GO:0005623, GO:0005737, GO:0005829, GO:0006464, GO:0006508, GO:0006515, GO:0006516 +33 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.456 · purifying
Polymorphic sites (≥ 0.1% of strains) 9 synonymous, 12 missense, 0 nonsense, 1 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 0.39% of strains (568) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Glyco_hydro_92NPF17678.8 6.1e-5644–274 Glycosyl hydrolase family 92 N-terminal domain
Glyco_hydro_92PF07971.19 1.9e-139280–749 Glycosyl hydrolase family 92 catalytic domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: lpqN (lipoprotein LpqN), high confidence from genomic context alone (score 772 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0583c lpqN lipoprotein LpqN 771 772 ctx neighborhood:771
Rv1733c transmembrane protein 451 451 ctx cooccurence:446
Rv3912 rsmA anti-sigma-M factor RsmA 407 408 ctx cooccurence:404
Rv3903c cpnT hyp hypothetical protein 873 64 textmining:870
Rv3551 CoA-transferase subunit alpha 575 52 textmining:571
Rv1030 kdpB potassium-transporting ATPase subunit B 662 47 textmining:660

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: glycosidase
  • MTBC0 PGAP product: GH92 family glycosyl hydrolase
  • Pfam (hmmscan --cut_ga): Glyco_hydro_92N PF17678.8 (E=6e-56), Glyco_hydro_92 PF07971.19 (E=2e-139)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215098.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Glyco_hydro_92N (PF17678.8), Glyco_hydro_92 (PF07971.19)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG3537
  • Curated reference: UniProt O86365 (SwissProt, reviewed; Inferred from homology)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 6 functional partner(s); context anchor lpqN
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000614|Rv0584|
MRARRLRRALAALLAVAGLFVPFIVGVPTAYDGEPVFVAIPVEHVNTLIGTGTGAAIVGEINNFPGASVPFGMVQYSPDTVDNYAGYDYDNPHSTGFSMTHASVGCPAFGDISMLPTTTPLGSQPWSAWEEIAHDDTEVGVPGYYTVRFPGTGVIAELTATTRTGVGRFRYPRNGWPALFHVRSGASLAGNYAATLQIEDNTTITGSATSGGFCGKKNLYTVYFAMKFSQPFSSYGTWDGYAVYPGSHSMNSSYSGGYVGFPAGSVLEVRTALSYVSVDGARANLDAEGGASFDDIRAATSSEWNAALSRIAVAGRGPGDVDTFYTCLYRSLLHPNTFNDVDGRYIGFDGVIHSVASGHTHYANFSDWDTYRSLAPLQGLLFPQRASDMIQSLVTDAEQSGAYPRWALANSATGMMSGDSVVPLIVNLYAFGARDFDLKSALHYMVNAATQGGVGLDGFLERPGIAAYLRLGYGPQTAEFRANGRIAGASVTLEWSVDDFAISRFADSLGDTATAAVFQNRSQYWQNLFNPTTGYISPRSAAGFFPDGPGFVAYPSGFGQDGYDEGNAEQYLWWVPHNVAGLVTALGGRTAVVKRLDRFTKKLNVGPNEPYLWAGNEPGFGVPWLYNYIGQPWKTQRTVDRVRGLFGPTPGGAPGNDDLGALSSWYVWAALGLYPSTPGTTILTVNTPLFDRAVIALPTGKSIQITAPGASGRNRLKYIDGLTIDRQPSNQTFLPESIVRTGGDLTFSLAGTPNKVWGTAASAAPPSFGAGSSAVTVNIARPIIGIVPGATGTVTVDAQRMIDGVDDYTVTPTSYVVGIAAEPLSGQFDDDGAVSASVAITVARSVPSGYYPIYVTTSAGDSARTLIVLVVVAEAVE