pks7 Resolved · high auto-curated

H37Rv Rv1661 · MTBC0 mtbc0_001770 · 2126 aa · 1887346–1893726 (+) · RefSeq NP_216177.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)polyketide synthase
MTBC0 PGAP re-annotationtype I polyketide synthase
Revised (this work)Type I polyketide synthase. Pfam: Docking (PF08990.17), ketoacyl-synt (PF00109.33), Ketoacyl-synt_C (PF02801.29), KAsynt_C_assoc (PF16197.12), CurL-like_PKS_C (PF22621.3), Acyl_transf_1 (PF00698.27), PKS_DH_N (PF21089.4), PS-DH (PF14765.13), SpnB_Rossmann (PF22953.4), ADH_N (PF08240.18), ADH_zinc_N (PF00107.33), ADH_zinc_N_2 (PF13602.13), KR (PF08659.17), adh_short (PF00106.32), Epimerase (PF01370.28), PP-binding (PF00550.32).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P94996 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable polyketide synthase Pks7

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category Q Secondary metabolites biosynthesis, transport and catabolism
Preferred namepks7
eggNOG descriptionpolyketide synthase
Orthologous groupCOG0604
KEGG orthology K12434

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.709 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 21 synonymous, 39 missense, 1 nonsense, 8 frameshift
Disruption 9 distinct premature-stop/frameshift site(s); most common in 5.94% of strains (8632) · convergent

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DockingPF08990.17 2.0e-064–31 Erythronolide synthase docking domain
ketoacyl-syntPF00109.33 1.4e-9535–284 Beta-ketoacyl synthase, N-terminal domain
Ketoacyl-synt_CPF02801.29 6.5e-42292–408 Beta-ketoacyl synthase, C-terminal domain
KAsynt_C_assocPF16197.12 2.2e-14411–528 Ketoacyl-synthetase C-terminal extension
CurL-like_PKS_CPF22621.3 8.3e-08479–539 CurL-like, PKS C-terminal
Acyl_transf_1PF00698.27 1.4e-97564–884 Acyl transferase domain
PKS_DH_NPF21089.4 2.3e-24934–1032 Polyketide synthase dehydratase domain
PS-DHPF14765.13 2.3e-241060–1204 Polyketide synthase dehydratase N-terminal domain
SpnB_RossmannPF22953.4 2.6e-341231–1352 Polyketide synthase extender module SpnB, Rossmann fold domain
ADH_NPF08240.18 1.0e-081395–1449 Alcohol dehydrogenase GroES-like domain
ADH_zinc_NPF00107.33 1.3e-111512–1601 Zinc-binding dehydrogenase
ADH_zinc_N_2PF13602.13 1.1e-181549–1679 Zinc-binding dehydrogenase
KRPF08659.17 3.7e-591694–1872 KR domain
adh_shortPF00106.32 2.8e-111695–1853 short chain dehydrogenase
EpimerasePF01370.28 3.0e-081696–1839 NAD dependent epimerase/dehydratase family
PP-bindingPF00550.32 5.2e-131982–2047 Phosphopantetheine attachment site

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: fas (fatty acid synthase), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2524c fas fatty acid synthase 999 1000 ctx neighborhood:544 coexpression:810 experimental:999 database:549 textmining:736
Rv2383c mbtB phenyloxazoline synthase 994 981 ctx neighborhood:544 cooccurence:419 coexpression:877 experimental:473 textmining:707
Rv1662 pks8 polyketide synthase 981 979 ctx neighborhood:748 coexpression:863 experimental:433
Rv1664 pks9 polyketide synthase 978 978 ctx neighborhood:855 coexpression:801
Rv1663 pks17 polyketide synthase 974 970 ctx neighborhood:749 coexpression:859
Rv2243 fabD malonyl CoA-acyl carrier protein transacylase 971 963 coexpression:602 experimental:787 database:549
Rv0101 nrp peptide synthetase Nrp 975 959 ctx cooccurence:498 coexpression:773 experimental:473 textmining:424
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 968 943 ctx neighborhood:544 experimental:795 textmining:470
Rv1527c pks5 polyketide synthase 966 943 ctx neighborhood:544 experimental:795 textmining:439
Rv2933 ppsC phthiocerol synthesis polyketide synthase type I PpsC 966 942 ctx neighborhood:544 experimental:795 textmining:439
Rv2940c mas multifunctional mycocerosic acid synthase 966 942 ctx neighborhood:544 experimental:795 textmining:439
Rv2048c pks12 polyketide synthase 958 930 ctx neighborhood:544 experimental:795 textmining:439
Rv3800c pks13 polyketide synthase 958 904 ctx neighborhood:544 coexpression:675 textmining:582
Rv0310c hyp hypothetical protein 897 864 coexpression:449 experimental:465 database:561
Rv3147 nuoC NADH-quinone oxidoreductase subunit C 885 862 coexpression:448 experimental:472 database:564

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: polyketide synthase
  • MTBC0 PGAP product: type I polyketide synthase
  • Pfam (hmmscan --cut_ga): Docking PF08990.17 (E=2e-06), ketoacyl-synt PF00109.33 (E=1e-95), Ketoacyl-synt_C PF02801.29 (E=6e-42), KAsynt_C_assoc PF16197.12 (E=2e-14), CurL-like_PKS_C PF22621.3 (E=8e-08), Acyl_transf_1 PF00698.27 (E=1e-97), PKS_DH_N PF21089.4 (E=2e-24), PS-DH PF14765.13 (E=2e-24), SpnB_Rossmann PF22953.4 (E=3e-34), ADH_N PF08240.18 (E=1e-08), ADH_zinc_N PF00107.33 (E=1e-11), ADH_zinc_N_2 PF13602.13 (E=1e-18), KR PF08659.17 (E=4e-59), adh_short PF00106.32 (E=3e-11), Epimerase PF01370.28 (E=3e-08), PP-binding PF00550.32 (E=5e-13)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216177.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Docking (PF08990.17), ketoacyl-synt (PF00109.33), Ketoacyl-synt_C (PF02801.29), KAsynt_C_assoc (PF16197.12), CurL-like_PKS_C (PF22621.3), Acyl_transf_1 (PF00698.27), PKS_DH_N (PF21089.4), PS-DH (PF14765.13), SpnB_Rossmann (PF22953.4), ADH_N (PF08240.18), ADH_zinc_N (PF00107.33), ADH_zinc_N_2 (PF13602.13), KR (PF08659.17), adh_short (PF00106.32), Epimerase (PF01370.28), PP-binding (PF00550.32)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0604
  • Curated reference: UniProt P94996 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 442 functional partner(s); context anchor fas
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001770|Rv1661|pks7
MNSTPEDLVKALRRSLKQNERLKRENRDLLARTTEPVAVVGMGCRYPGGVDSPETLWELVAHGRDAVSEFPADRGWDVAGLFDPDPDAVGKSYTRCGGFLTDVAGFDAEFFGIAPSEALAMDPQQRLLLEVSWEALERAGIDPITLRGSQTGVFAGVFHGSYGGQGRVPGDLERYGLRGSTLSVASGRVAYVLGLQGPAVSVDTACSSSLVALHLAVQSLRLGECDLALVGGVTVMATPAMFIEFSRQRALSADGRCKAYAGAADGTAFAEGAGVLVLARLADARRLGHPVLALVRGSAVNQDGASNGLATPNGPAQQRVITAALASARLGVADVDVVEGHGTGTTLGDPIEAQAILATYGQRPADRPLWLGSIKSNIGHTSAAAGVAGVIKMVQAMRHGVLPKTLHVDVPTPHVDWSAGAVSLLTEPRPWHVPGRPRRAGVSSFGISGTNAHVILEEAPAVEPVGAAHGNDPVAVPWVLSARSAQALTNQARRLLAWVGADENVRPLDVGWSLVNTRSLFDHRAVVVGADRTQLMEGLTGLAAGVPGADVVAGRAQTVGKTAFVFPGQGAQWLGMGAQLCATAPVFAEHIHRCERALREHVEWSLLDVLRGAPGAPGLDRVDVVQPALWAVMVSLAELWRSVGVVPDAVIGHSQGEIAAAYVAGALSLRDAAAVVALRSRLLVRLGGAGGMVSLACGQPQAEKLASQWGDRLNIAAVNGVSSVVLAGETDAVTELMQRCEAEGIRARRIDVDYASHSAQVDAIREELIAALRGIEPRTSTVAFFSTVTGELMDTAGVNAEYWYRSIRQPVQFERAVRNAFDGGYRVFVESSPHPVLIAGIEETLVDCDRGATGEPIVIPTLGRDDGGVGRFWLSAGQAHVAGVGVDWRAAFADLGGRRVELPTYAFARQRFWLDGLGAVGGDLGGVGLVGAEHGLLAAVVQRPDSGGVVLTGRISVVAAPWLADHAVGPVVLFPGTGFVELALRAGDEVGCSVLQELTLQAPLVLPADGVRVQVVVGGVEQSGTRNVWVYSAAGQADSSPGWTLHAQGVLGVGSVQPAAELSVWPPVGARAMDVADGYQVLAARGYGYGPAFRGLQALWRRGAEVFADVTLPEGVPIRGFGIHPAVLDAALHAWGIVEGEQQTMLPFSWQGVCLHASGAARVRVRLAPVGRGAVSVELADPQGLPVLSVRQLMVRPVSAAALSRSTAGDRGLLEMIWTPVPLEGGDIGDDAVVWELPPHAGAQAGGDVLAAVYRGVHEVLEVLQSWLASDATGLGVVVTRGAVGPVDDDVTDLAGAAVWGLVRSAQAEHPGRVVLVDTDGSVAVEDAVGFGARSGEPQLVVRRGRVYAARLAPVAAGLTLPSASAGGWRLVAGGGGTLADVVVAPVAPVELATGQVRVAVGAVGVNFRDVLVALGMYPGGGELGVDGAGVVVEVGPGVTGLAVGDRVMGLLGLVGSEAVVDARLVTMVPAGWSLVEAAAVPVAFLTAFYGLSVLAEVAAGQKVLVHAGTGGVGMAAVSLARYWGAEVFVTASRAKWDTLRAMGFDDIHISDSRSLEFEEAFLRATEGSGVDVVLNSLAGEFTDASLRLLPSGGRFIELGKTDIRDGQTVAERHRGVRYRAFDLVEAGPDRIAAMLSEVVGLLAAGVLARLPVKTFDARCAPAAYRFVSQARHIGKVVLTIPDGPGGQSGLAGGTVVVTGGTGMAGSAVATHLVRRHGVANLVLVSRSGEQADRAAEVAALLREGGAQVAVVSCDVADRDALAALLAGLDPRYPLKGVFHAAGVLDDAVITGLTPDRVDTVLRAKVDGAWNLHELTEDMDLSAFVVFSSMAGIVGTPAQGNYAAANAFLDGLVAYRRSRGLAGLSVAWGLWEQASAMTRHLGERDRARMTQAGLAPLTTEQALGFLDTALQADRAVVVAARLDRAALAGAGAALPALFSQLAAGPTRRRIDAADTAVSMSGLVSRLHALTPERRQRELTDLVISNAAAVLGRSSSVDINAHKAFQDLGFDSLTAVELRNRLKTATGLTLSPTLIFDYPTPATLAEHLDSRLVTASGSDQQSLSDRVDDITRELVVLLDQPDLSANVKAHLRTRLQTMLTSLTTEDDDIAAATESQLFAILDEELGS