cyp132 Resolved · high auto-curated

H37Rv Rv1394c · MTBC0 - · 461 aa · 1569584–1570969 (-) · RefSeq YP_177807.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)cytochrome P450 Cyp132
MTBC0 PGAP re-annotation
Revised (this work)Cytochrome P450 Cyp132. Pfam: p450 (PF00067.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Annotated on the H37Rv protein: this gene has no 1:1 ancestral MTBC0 anchor (PE/PPE, paralogue, IS element, or otherwise unanchored CDS).

Curated reference (UniProt)

UniProt P9WPN3 SwissProt · reviewed · Evidence at protein level
UniProt namePutative cytochrome P450 132
EC (curated) EC 1.14.-.-

UniProt still lists this protein as Putative cytochrome P450 132; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category C Energy production and conversion
Preferred namecyp132
eggNOG descriptionCytochrome P450
Orthologous groupCOG2124

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.333 · purifying
Polymorphic sites (≥ 0.1% of strains) 8 synonymous, 7 missense, 1 nonsense, 0 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 0.20% of strains (285) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
p450PF00067.28 1.2e-6445–443 Cytochrome P450

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1393c (monoxygenase), high confidence from genomic context alone (score 985 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1393c monoxygenase 985 985 ctx neighborhood:882 coexpression:860
Rv3800c pks13 polyketide synthase 943 938 experimental:891
Rv2380c mbtE peptide synthetase 809 800 experimental:689
Rv1937 oxygenase 812 785 experimental:478
Rv1395 HTH-type transcriptional regulator 862 767 ctx neighborhood:764 textmining:433
Rv3554 fdxB electron transfer protein FdxB 742 710
Rv2268c cyp128 cytochrome P450 Cyp128 860 703 ctx cooccurence:700 textmining:548
Rv2932 ppsB phthiocerol synthesis polyketide synthase type I PpsB 719 701 experimental:460
Rv0766c cyp123 cytochrome P450 Cyp123 795 689 ctx cooccurence:685
Rv0719 rplF 50S ribosomal protein L6 689 689 experimental:412 database:493
Rv1629 polA DNA polymerase I 706 688 database:638
Rv3571 kshB 3-ketosteroid-9-alpha-hydroxylase reductase subunit 703 686
Rv2946c pks1 polyketide synthase 716 682 experimental:460
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 699 666
Rv1785c cyp143 cytochrome P450 Cyp143 831 664 ctx cooccurence:662 textmining:518

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Annotation from H37Rv (no MTBC0 1:1 anchor; H37Rv protein used): cytochrome P450 Cyp132
  • Pfam (hmmscan --cut_ga): p450 PF00067.28 (E=1e-64)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq YP_177807.1)
  • Domains: Pfam-A via hmmscan --cut_ga — p450 (PF00067.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2124
  • Curated reference: UniProt P9WPN3 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 123 functional partner(s); context anchor Rv1393c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>H37Rv|Rv1394c|cyp132
MATATTQRPLKGPAKRMSTWTMTREAITIGFDAGDGFLGRLRGSDITRFRCAGRRFVSISHPDYVDHVLHEARLKYVKSDEYGPIRATAGLNLLTDEGDSWARHRGALNSTFARRHLRGLVGLMIDPIADVTAARVPGAQFDMHQSMVETTLRVVANALFSQDFGPLVQSMHDLATRGLRRAEKLERLGLWGLMPRTVYDTLIWCIYSGVHLPPPLREMQEITLTLDRAINSVIDRRLAEPTNSADLLNVLLSADGGIWPRQRVRDEALTFMLAGHETTANAMSWFWYLMALNPQARDHMLTELDDVLGMRRPTADDLGKLAWTTACLQESQRYFSSVWIIAREAVDDDIIDGHRIRRGTTVVIPIHHIHHDPRWWPDPDRFDPGRFLRCPTDRPRCAYLPFGGGRRICIGQSFALMEMVLMAAIMSQHFTFDLAPGYHVELEATLTLRPKHGVHVIGRRR