Rv3200c Family assigned · medium auto-curated

H37Rv Rv3200c · MTBC0 mtbc0_003404 · 355 aa · 3594744–3595811 (-) · RefSeq NP_217716.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane cation transporter
MTBC0 PGAP re-annotationpotassium channel family protein
Revised (this work)Potassium channel family protein. Pfam: Ion_trans_2 (PF07885.23), TrkA_N (PF02254.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53346 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible transmembrane cation transporter

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category P Inorganic ion transport and metabolism
Preferred namekch
eggNOG descriptiontransport
Orthologous groupCOG1226
KEGG orthology K10716

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.654 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 4 synonymous, 7 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Ion_trans_2PF07885.23 2.4e-1448–129 Ion channel
TrkA_NPF02254.25 2.6e-27150–266 TrkA-N domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: nudC (NADH pyrophosphatase), high confidence from genomic context alone (score 810 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3199c nudC NADH pyrophosphatase 810 810 ctx neighborhood:804
Rv3202c adnA ATP-dependent DNA helicase 813 806 ctx neighborhood:801
Rv3201c adnB ATP-dependent DNA helicase 812 804 ctx neighborhood:801
Rv1407 fmu 16S rRNA m5C967 methyltransferase 675 675 coexpression:578
Rv0511 hemD uroporphyrin-III C-methyltransferase 553 554 coexpression:454
Rv0948c chorismate mutase 544 544 ctx cooccurence:531
Rv3013 hyp hypothetical protein 478 478 ctx cooccurence:460
Rv0260c transcriptional regulator 470 470 coexpression:452
Rv3119 moaE1 molybdopterin synthase catalytic subunit 1 465 466 coexpression:465
Rv3323c moaX MoaD-MoaE fusion protein MoaX 464 463 coexpression:458
Rv0813c hyp hypothetical protein 462 463 ctx cooccurence:461
Rv0866 moaE2 molybdopterin synthase catalytic subunit 2 461 462 coexpression:461
Rv1631 coaE dephospho-CoA kinase CoaE 454 454 coexpression:423
Rv3753c hyp hypothetical protein 421 422 ctx cooccurence:420
Rv3311 hyp hypothetical protein 411 411 ctx cooccurence:409

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane cation transporter
  • MTBC0 PGAP product: potassium channel family protein
  • Pfam (hmmscan --cut_ga): Ion_trans_2 PF07885.23 (E=2e-14), TrkA_N PF02254.25 (E=3e-27)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217716.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Ion_trans_2 (PF07885.23), TrkA_N (PF02254.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1226
  • Curated reference: UniProt O53346 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 30 functional partner(s); context anchor nudC
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003404|Rv3200c|
MAGSWRRLRGLDEKLTAQPGYALVGVLRIPQRRASPARVISRRVVVAVVALLLTAGIVYVDRDGYLDAQGDRLTFLDCLYYAAVTLSTTGYGDITPISEFARAINIFVITPLRIAFLILLVGTTLEVLTETSRQAYKIQRWRSRVRNHTVVIGYGTKGKTAVAAMVSDELVPGEIVVVDTDSGVLERAAAAGLVTVHGDATKSDVLRLAGTQHASSIIVATSRDDTAVLVTLTAREIAPKAKIVASIREAENQHLLRQSGADTVVVSSETAGRLLGIATTTPSVVEMIEDLLTPEAGLAVAEREVEQAEVGGSPRHLRDIVLGVVRDGQLLRIGAPEVDAIEASDRLLYIRQVGR