Rv3190c Still unknown · low auto-curated

H37Rv Rv3190c · MTBC0 mtbc0_003387 · 421 aa · 3577554–3578819 (-) · RefSeq NP_217706.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationhypothetical protein
Revised (this work)Conserved hypothetical protein; no recognised domain. Function unknown.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53336 TrEMBL · unreviewed · Evidence at protein level
UniProt nameRestriction endonuclease

Functional vocabulary (eggNOG-mapper, orthology transfer)

Orthologous grouparCOG12798

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 1.095 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 5 missense, 2 nonsense, 0 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 5.95% of strains (8635) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: PPE8 (PPE family protein PPE8), high confidence from genomic context alone (score 730 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv0355c PPE8 PPE family protein PPE8 729 730 ctx cooccurence:727
Rv1452c PE_PGRS28 PE-PGRS family protein PE_PGRS28 728 729 ctx cooccurence:722
Rv3347c PPE55 PPE family protein PPE55 725 726 ctx cooccurence:724
Rv3350c PPE56 PPE family protein PPE56 723 723 ctx cooccurence:722
Rv2209 integral membrane protein 716 716 ctx cooccurence:716
Rv2490c PE_PGRS43 PE-PGRS family protein PE_PGRS43 714 714 ctx cooccurence:707
Rv1004c membrane protein 713 713 ctx cooccurence:713
Rv1651c PE_PGRS30 PE-PGRS family protein PE_PGRS30 706 706 ctx cooccurence:699
Rv0304c PPE5 PPE family protein PPE5 700 701 ctx cooccurence:699
Rv1917c PPE34 PPE family protein PPE34 698 699 ctx cooccurence:698
Rv3343c PPE54 PPE family protein PPE54 690 690 ctx cooccurence:687
Rv2082 hyp hypothetical protein 687 688 ctx cooccurence:677
Rv1045 hyp hypothetical protein 684 684 ctx cooccurence:683
Rv0872c PE_PGRS15 PE-PGRS family protein PE_PGRS15 681 682 ctx cooccurence:674
Rv3879c espK ESX-1 secretion-associated protein EspK 674 675 ctx cooccurence:667

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: hypothetical protein
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217706.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG arCOG12798
  • Curated reference: UniProt O53336 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 64 functional partner(s); context anchor PPE8
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003387|Rv3190c|
MEYVQLFSKGRLNDLAGSLAGFLGKASQATAQRLQSWDADDLLNTPVDDVVEQLVELGSVECPDLRVDDAFMLPATEVDQQYRDWGEQRTRRVTRLVLVVPFEGHKDIFNLRPDQFTTMPPQVLRLQGHEIHLAIDNPSNDAAAINAAFHKQIANIEKYLGWSRRQIDLHNQGLRNELPGMVARRREQLLATRNLQAEIGFPVRRRKDADTYAAPISRKSVRPRPHRPAGARAAFKPEPAMQDEDYQSALRVLRNQRNALERTPSVAAKLDGEEIRDMLLVGLNAQFEGDAGGELFNGAGKTDILIRVDDRNIFIGECKVWSGPRTMDDALKQLFGYLVWRDTKAAILLFIRNKDVTAVIDNAIAKIKEHPNHKRCPAHRAGADQYEFTMHADGDPEREIHLTLIPFALRPTAEVPTTTIP